Now becoming the device that can restore Earth's environment, the hope-carrying Yamato is on its way back to Earth. They encounter the powerless vessel containing Cerestella, the last of the Jirels and is a part of Desler's cabinet. Along with General Goer and his fleet, which was dispatched by Captain Flakken and his crew (now including the ex-Yamato engine room operator Sukeji Yabu) using the dimensional submarine, Desler, who was presumed dead, comes at the Yamato for a final confrontation at the hyperspace gate...